WisdomTree Investments, the world’s most-shorted asset manager, racked up the biggest exchange-traded-fund outflows of the top 20 providers during Q1 2016, with $5.2 billion leaving the firm during the first three months of the year.

WisdomTree saw $1.5 billion leave its funds in March, tipping total outflows for the first quarter past the $5 billion mark, according to the data provider ETFGI. The outflows marked a reversal in fortunes from a year ago.